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CHAPTER
III. STRUGGLE FOR EXISTENCE. Bears on natural selection—The term used in a wide sense—Geometrical powers of increase—Rapid increase of naturalised animals and plants—Nature of the checks to increase—Competition universal—Effects of climate—Protection from the number of individuals—Complex relations of all animals and plants throughout nature—Struggle for life most severe between individuals and varieties of the same species; often severe between species of the same genus—The relation of organism to organism the most important of all relations. BEFORE
entering on the subject of this chapter, I must make a few preliminary remarks,
to show how the struggle for existence bears on Natural Selection. It has been
seen in the last chapter that amongst organic beings in a state of nature there
is some individual variability; indeed I am not aware that this has ever been
disputed. It is immaterial for us whether a multitude of doubtful forms be
called species or sub-species or varieties; what rank, for instance, the two or
three hundred doubtful forms of British plants are entitled to hold, if the
existence of any well-marked varieties be admitted. But the mere existence of
individual variability and of some few well-marked varieties, though necessary
as the foundation for the work, helps us but little in understanding how
species arise in nature. How have all those exquisite adaptations of one part
of the organisation to another part, and to the conditions of life, and of one
distinct organic being to another being, been perfected? We see these beautiful
co-adaptations most plainly in the woodpecker and missletoe; and only a little
less plainly in the humblest parasite which clings to the hairs of a quadruped
or feathers of a bird; in the structure of the beetle which dives through the
water; in the plumed seed which is wafted by the gentlest breeze; in short, we
see beautiful adaptations everywhere and in every part of the organic world. Again, it
may be asked, how is it that varieties, which I have called incipient species,
become ultimately converted into good and distinct species, which in most cases
obviously differ from each other far more than do the varieties of the same
species? How do those groups of species, which constitute what are called
distinct genera, and which differ from each other more than do the species of
the same genus, arise? All these results, as we shall more fully see in the
next chapter, follow inevitably from the struggle for life. Owing to this
struggle for life, any variation, however slight and from whatever cause
proceeding, if it be in any degree profitable to an individual of any species, in
its infinitely complex relations to other organic beings and to external
nature, will tend to the preservation of that individual, and will generally be
inherited by its offspring. The offspring, also, will thus have a better chance
of surviving, for, of the many individuals of any species which are
periodically born, but a small number can survive. I have called this
principle, by which each slight variation, if useful, is preserved, by the term
of Natural Selection, in order to mark its relation to man’s power of
selection. We have seen that man by selection can certainly produce great
results, and can adapt organic beings to his own uses, through the accumulation
of slight but useful variations, given to him by the hand of Nature. But
Natural Selection, as we shall hereafter see, is a power incessantly ready for
action, and is as immeasurably superior to man’s feeble efforts, as the works
of Nature are to those of Art. We will
now discuss in a little more detail the struggle for existence. In my future work
this subject shall be treated, as it well deserves, at much greater length. The
elder De Candolle and Lyell have largely and philosophically shown that all
organic beings are exposed to severe competition. In regard to plants, no one
has treated this subject with more spirit and ability than W. Herbert, Dean of
Manchester, evidently the result of his great horticultural knowledge. Nothing
is easier than to admit in words the truth of the universal struggle for life,
or more difficult—at least I have found it so—than constantly to bear this
conclusion in mind. Yet unless it be thoroughly engrained in the mind, I am
convinced that the whole economy of nature, with every fact on distribution,
rarity, abundance, extinction, and variation, will be dimly seen or quite
misunderstood. We behold the face of nature bright with gladness, we often see
superabundance of food; we do not see, or we forget, that the birds which are
idly singing round us mostly live on insects or seeds, and are thus constantly
destroying life; or we forget how largely these songsters, or their eggs, or
their nestlings, are destroyed by birds and beasts of prey; we do not always
bear in mind, that though food may be now superabundant, it is not so at all
seasons of each recurring year. I should
premise that I use the term Struggle for Existence in a large and metaphorical
sense, including dependence of one being on another, and including (which is
more important) not only the life of the individual, but success in leaving
progeny. Two canine animals in a time of dearth, may be truly said to struggle
with each other which shall get food and live. But a plant on the edge of a
desert is said to struggle for life against the drought, though more properly
it should be said to be dependent on the moisture. A plant
which annually produces a thousand seeds, of which on an average only one comes
to maturity, may be more truly said to struggle with the plants of the same and
other kinds which already clothe the ground. The missletoe is dependent on the
apple and a few other trees, but can only in a far-fetched sense be said to
struggle with these trees, for if too many of these parasites grow on the same
tree, it will languish and die. But several seedling missletoes, growing close
together on the same branch, may more truly be said to struggle with each
other. As the missletoe is disseminated by birds, its existence depends on
birds; and it may metaphorically be said to struggle with other fruit-bearing
plants, in order to tempt birds to devour and thus disseminate its seeds rather
than those of other plants. In these several senses, which pass into each
other, I use for convenience sake the general term of struggle for existence. A struggle
for existence inevitably follows from the high rate at which all organic beings
tend to increase. Every being, which during its natural lifetime produces
several eggs or seeds, must suffer destruction during some period of its life,
and during some season or occasional year, otherwise, on the principle of
geometrical increase, its numbers would quickly become so inordinately great
that no country could support the product. Hence, as more individuals are
produced than can possibly survive, there must in every case be a struggle for
existence, either one individual with another of the same species, or with the
individuals of distinct species, or with the physical conditions of life. It is
the doctrine of Malthus applied with manifold force to the whole animal and
vegetable kingdoms; for in this case there can be no artificial increase of
food, and no prudential restraint from marriage. Although some species may be
now increasing, more or less rapidly, in numbers, all cannot do so, for the
world would not hold them. There is
no exception to the rule that every organic being naturally increases at so
high a rate, that if not destroyed, the earth would soon be covered by the
progeny of a single pair. Even slow-breeding man has doubled in twenty-five
years, and at this rate, in a few thousand years, there would literally not be
standing room for his progeny. Linnæus has calculated that if an annual plant
produced only two seeds—and there is no plant so unproductive as this—and their
seedlings next year produced two, and so on, then in twenty years there would
be a million plants. The elephant is reckoned to be the slowest breeder of all
known animals, and I have taken some pains to estimate its probable minimum
rate of natural increase: it will be under the mark to assume that it breeds
when thirty years old, and goes on breeding till ninety years old, bringing
forth three pair of young in this interval; if this be so, at the end of the
fifth century there would be alive fifteen million elephants, descended from
the first pair. But we
have better evidence on this subject than mere theoretical calculations,
namely, the numerous recorded cases of the astonishingly rapid increase of
various animals in a state of nature, when circumstances have been favourable
to them during two or three following seasons. Still more striking is the evidence
from our domestic animals of many kinds which have run wild in several parts of
the world: if the statements of the rate of increase of slow-breeding cattle
and horses in South-America, and latterly in Australia, had not been well
authenticated, they would have been quite incredible. So it is with plants:
cases could be given of introduced plants which have become common throughout
whole islands in a period of less than ten years. Several of the plants now
most numerous over the wide plains of La Plata, clothing square leagues of
surface almost to the exclusion of all other plants, have been introduced from
Europe; and there are plants which now range in India, as I hear from Dr.
Falconer, from Cape Comorin to the Himalaya, which have been imported from
America since its discovery. In such cases, and endless instances could be
given, no one supposes that the fertility of these animals or plants has been
suddenly and temporarily increased in any sensible degree. The obvious
explanation is that the conditions of life have been very favourable, and that
there has consequently been less destruction of the old and young, and that
nearly all the young have been enabled to breed. In such cases the geometrical
ratio of increase, the result of which never fails to be surprising, simply
explains the extraordinarily rapid increase and wide diffusion of naturalised
productions in their new homes. In a state
of nature almost every plant produces seed, and amongst animals there are very
few which do not annually pair. Hence we may confidently assert, that all
plants and animals are tending to increase at a geometrical ratio, that all
would most rapidly stock every station in which they could any how exist, and
that the geometrical tendency to increase must be checked by destruction at
some period of life. Our familiarity with the larger domestic animals tends, I
think, to mislead us: we see no great destruction falling on them, and we
forget that thousands are annually slaughtered for food, and that in a state of
nature an equal number would have somehow to be disposed of. The only
difference between organisms which annually produce eggs or seeds by the
thousand, and those which produce extremely few, is, that the slow-breeders
would require a few more years to people, under favourable conditions, a whole
district, let it be ever so large. The condor lays a couple of eggs and the
ostrich a score, and yet in the same country the condor may be the more
numerous of the two: the Fulmar petrel lays but one egg, yet it is believed to
be the most numerous bird in the world. One fly deposits hundreds of eggs, and
another, like the hippobosca, a single one; but this difference does not
determine how many individuals of the two species can be supported in a
district. A large number of eggs is of some importance to those species, which
depend on a rapidly fluctuating amount of food, for it allows them rapidly to
increase in number. But the real importance of a large number of eggs or seeds
is to make up for much destruction at some period of life; and this period in
the great majority of cases is an early one. If an animal can in any way
protect its own eggs or young, a small number may be produced, and yet the
average stock be fully kept up; but if many eggs or young are destroyed, many
must be produced, or the species will become extinct. It would suffice to keep
up the full number of a tree, which lived on an average for a thousand years,
if a single seed were produced once in a thousand years, supposing that this
seed were never destroyed, and could be ensured to germinate in a fitting
place. So that in all cases, the average number of any animal or plant depends
only indirectly on the number of its eggs or seeds. In looking
at Nature, it is most necessary to keep the foregoing considerations always in
mind—never to forget that every single organic being around us may be said to
be striving to the utmost to increase in numbers; that each lives by a struggle
at some period of its life; that heavy destruction inevitably falls either on
the young or old, during each generation or at recurrent intervals. Lighten any
check, mitigate the destruction ever so little, and the number of the species
will almost instantaneously increase to any amount. The face of Nature may be
compared to a yielding surface, with ten thousand sharp wedges packed close
together and driven inwards by incessant blows, sometimes one wedge being
struck, and then another with greater force. What
checks the natural tendency of each species to increase in number is most obscure.
Look at the most vigorous species; by as much as it swarms in numbers, by so
much will its tendency to increase be still further increased. We know not
exactly what the checks are in even one single instance. Nor will this surprise
any one who reflects how ignorant we are on this head, even in regard to
mankind, so incomparably better known than any other animal. This subject has
been ably treated by several authors, and I shall, in my future work, discuss
some of the checks at considerable length, more especially in regard to the
feral animals of South America. Here I will make only a few remarks, just to
recall to the reader’s mind some of the chief points. Eggs or very young
animals seem generally to suffer most, but this is not invariably the case.
With plants there is a vast destruction of seeds, but, from some observations
which I have made, I believe that it is the seedlings which suffer most from
germinating in ground already thickly stocked with other plants. Seedlings,
also, are destroyed in vast numbers by various enemies; for instance, on a
piece of ground three feet long and two wide, dug and cleared, and where there
could be no choking from other plants, I marked all the seedlings of our native
weeds as they came up, and out of the 357 no less than 295 were destroyed,
chiefly by slugs and insects. If turf which has long been mown, and the case
would be the same with turf closely browsed by quadrupeds, be let to grow, the
more vigorous plants gradually kill the less vigorous, though fully grown,
plants: thus out of twenty species growing on a little plot of turf (three feet
by four) nine species perished from the other species being allowed to grow up
freely. The amount
of food for each species of course gives the extreme limit to which each can
increase; but very frequently it is not the obtaining food, but the serving as
prey to other animals, which determines the average numbers of a species. Thus,
there seems to be little doubt that the stock of partridges, grouse, and hares
on any large estate depends chiefly on the destruction of vermin. If not one
head of game were shot during the next twenty years in England, and, at the
same time, if no vermin were destroyed, there would, in all probability, be
less game than at present, although hundreds of thousands of game animals are
now annually killed. On the other hand, in some cases, as with the elephant and
rhinoceros, none are destroyed by beasts of prey: even the tiger in India most
rarely dares to attack a young elephant protected by its dam. Climate
plays an important part in determining the average numbers of a species, and
periodical seasons of extreme cold or drought, I believe to be the most
effective of all checks. I estimated that the winter of 1854-55 destroyed
four-fifths of the birds in my own grounds; and this is a tremendous
destruction, when we remember that ten per cent. is an extraordinarily severe
mortality from epidemics with man. The action of climate seems at first sight
to be quite independent of the struggle for existence; but in so far as climate
chiefly acts in reducing food, it brings on the most severe struggle between
the individuals, whether of the same or of distinct species, which subsist on
the same kind of food. Even when climate, for instance extreme cold, acts
directly, it will be the least vigorous, or those which have got least food
through the advancing winter, which will suffer most. When we travel from south
to north, or from a damp region to a dry, we invariably see some species
gradually getting rarer and rarer, and finally disappearing; and the change of
climate being conspicuous, we are tempted to attribute the whole effect to its
direct action. But this is a very false view: we forget that each species, even
where it most abounds, is constantly suffering enormous destruction at some
period of its life, from enemies or from competitors for the same place and
food; and if these enemies or competitors be in the least degree favoured by
any slight change of climate, they will increase in numbers, and, as each area
is already fully stocked with inhabitants, the other species will decrease.
When we travel southward and see a species decreasing in numbers, we may feel
sure that the cause lies quite as much in other species being favoured, as in
this one being hurt. So it is when we travel northward, but in a somewhat
lesser degree, for the number of species of all kinds, and therefore of
competitors, decreases northwards; hence in going northward, or in ascending a
mountain, we far oftener meet with stunted forms, due to the directly
injurious action of climate, than we do in proceeding southwards or in
descending a mountain. When we reach the Arctic regions, or snow-capped
summits, or absolute deserts, the struggle for life is almost exclusively with
the elements. That
climate acts in main part indirectly by favouring other species, we may clearly
see in the prodigious number of plants in our gardens which can perfectly well
endure our climate, but which never become naturalised, for they cannot compete
with our native plants, nor resist destruction by our native animals. When a
species, owing to highly favourable circumstances, increases inordinately in
numbers in a small tract, epidemics—at least, this seems generally to occur
with our game animals—often ensue: and here we have a limiting check
independent of the struggle for life. But even some of these so-called
epidemics appear to be due to parasitic worms, which have from some cause,
possibly in part through facility of diffusion amongst the crowded animals, been
disproportionably favoured: and here comes in a sort of struggle between the
parasite and its prey. On the
other hand, in many cases, a large stock of individuals of the same species,
relatively to the numbers of its enemies, is absolutely necessary for its
preservation. Thus we can easily raise plenty of corn and rape-seed, &c.,
in our fields, because the seeds are in great excess compared with the number
of birds which feed on them; nor can the birds, though having a superabundance
of food at this one season, increase in number proportionally to the supply of
seed, as their numbers are checked during winter: but any one who has tried,
knows how troublesome it is to get seed from a few wheat or other such plants
in a garden; I have in this case lost every single seed. This view of the
necessity of a large stock of the same species for its preservation, explains,
I believe, some singular facts in nature, such as that of very rare plants
being sometimes extremely abundant in the few spots where they do occur; and
that of some social plants being social, that is, abounding in individuals,
even on the extreme confines of their range. For in such cases, we may believe,
that a plant could exist only where the conditions of its life were so
favourable that many could exist together, and thus save each other from utter
destruction. I should add that the good effects of frequent intercrossing, and
the ill effects of close interbreeding, probably come into play in some of
these cases; but on this intricate subject I will not here enlarge. Many cases
are on record showing how complex and unexpected are the checks and relations
between organic beings, which have to struggle together in the same country. I
will give only a single instance, which, though a simple one, has interested
me. In Staffordshire, on the estate of a relation where I had ample means of
investigation, there was a large and extremely barren heath, which had never
been touched by the hand of man; but several hundred acres of exactly the same
nature had been enclosed twenty-five years previously and planted with Scotch
fir. The change in the native vegetation of the planted part of the heath was
most remarkable, more than is generally seen in passing from one quite
different soil to another: not only the proportional numbers of the
heath-plants were wholly changed, but twelve species of plants (not counting
grasses and carices) flourished in the plantations, which could not be found on
the heath. The effect on the insects must have been still greater, for six
insectivorous birds were very common in the plantations, which were not to be
seen on the heath; and the heath was frequented by two or three distinct
insectivorous birds. Here we see how potent has been the effect of the
introduction of a single tree, nothing whatever else having been done, with the
exception that the land had been enclosed, so that cattle could not enter. But
how important an element enclosure is, I plainly saw near Farnham, in Surrey.
Here there are extensive heaths, with a few clumps of old Scotch firs on the
distant hill-tops: within the last ten years large spaces have been enclosed,
and self-sown firs are now springing up in multitudes, so close together that
all cannot live. When I
ascertained that these young trees had not been sown or planted, I was so much
surprised at their numbers that I went to several points of view, whence I
could examine hundreds of acres of the unenclosed heath, and literally I could
not see a single Scotch fir, except the old planted clumps. But on looking
closely between the stems of the heath, I found a multitude of seedlings and
little trees, which had been perpetually browsed down by the cattle. In one
square yard, at a point some hundred yards distant from one of the old clumps,
I counted thirty-two little trees; and one of them, judging from the rings of
growth, had during twenty-six years tried to raise its head above the stems of
the heath, and had failed. No wonder that, as soon as the land was enclosed, it
became thickly clothed with vigorously growing young firs. Yet the heath was so
extremely barren and so extensive that no one would ever have imagined that
cattle would have so closely and effectually searched it for food. Here we
see that cattle absolutely determine the existence of the Scotch fir; but in
several parts of the world insects determine the existence of cattle. Perhaps
Paraguay offers the most curious instance of this; for here neither cattle nor
horses nor dogs have ever run wild, though they swarm southward and northward
in a feral state; and Azara and Rengger have shown that this is caused by the
greater number in Paraguay of a certain fly, which lays its eggs in the navels
of these animals when first born. The increase of these flies, numerous as they
are, must be habitually checked by some means, probably by birds. Hence, if
certain insectivorous birds (whose numbers are probably regulated by hawks or
beasts of prey) were to increase in Paraguay, the flies would decrease—then
cattle and horses would become feral, and this would certainly greatly alter
(as indeed I have observed in parts of South America) the vegetation: this
again would largely affect the insects; and this, as we just have seen in
Staffordshire, the insectivorous birds, and so onwards in ever-increasing circles
of complexity. We began this series by insectivorous birds, and we have ended
with them. Not that in nature the relations can ever be as simple as this.
Battle within battle must ever be recurring with varying success; and yet in
the long-run the forces are so nicely balanced, that the face of nature remains
uniform for long periods of time, though assuredly the merest trifle would
often give the victory to one organic being over another. Nevertheless so
profound is our ignorance, and so high our presumption, that we marvel when we
hear of the extinction of an organic being; and as we do not see the cause, we
invoke cataclysms to desolate the world, or invent laws on the duration of the
forms of life! I am
tempted to give one more instance showing how plants and animals, most remote
in the scale of nature, are bound together by a web of complex relations. I
shall hereafter have occasion to show that the exotic Lobelia fulgens, in this
part of England, is never visited by insects, and consequently, from its peculiar
structure, never can set a seed. Many of our orchidaceous plants absolutely
require the visits of moths to remove their pollen-masses and thus to fertilise
them. I have, also, reason to believe that humble-bees are indispensable to the
fertilisation of the heartsease (Viola tricolor), for other bees do not visit
this flower. From experiments which I have tried, I have found that the visits
of bees, if not indispensable, are at least highly beneficial to the
fertilisation of our clovers; but humble-bees alone visit the common red clover
(Trifolium pratense), as other bees cannot reach the nectar. Hence I have very
little doubt, that if the whole genus of humble-bees became extinct or very
rare in England, the heartsease and red clover would become very rare, or
wholly disappear. The number of humble-bees in any district depends in a great
degree on the number of field-mice, which destroy their combs and nests; and
Mr. H. Newman, who has long attended to the habits of humble-bees, believes
that “more than two thirds of them are thus destroyed all over England.” Now
the number of mice is largely dependent, as every one knows, on the number of
cats; and Mr. Newman says, “Near villages and small towns I have found the
nests of humble-bees more numerous than elsewhere, which I attribute to the
number of cats that destroy the mice.” Hence it is quite credible that the
presence of a feline animal in large numbers in a district might determine,
through the intervention first of mice and then of bees, the frequency of
certain flowers in that district! In the
case of every species, many different checks, acting at different periods of
life, and during different seasons or years, probably come into play; some one
check or some few being generally the most potent, but all concurring in
determining the average number or even the existence of the species. In some
cases it can be shown that widely-different checks act on the same species in
different districts. When we look at the plants and bushes clothing an entangled
bank, we are tempted to attribute their proportional numbers and kinds to what
we call chance. But how false a view is this! Every one has heard that when an
American forest is cut down, a very different vegetation springs up; but it has
been observed that the trees now growing on the ancient Indian mounds, in the
Southern United States, display the same beautiful diversity and proportion of
kinds as in the surrounding virgin forests. What a struggle between the several
kinds of trees must here have gone on during long centuries, each annually
scattering its seeds by the thousand; what war between insect and
insect—between insects, snails, and other animals with birds and beasts of
prey—all striving to increase, and all feeding on each other or on the trees or
their seeds and seedlings, or on the other plants which first clothed the
ground and thus checked the growth of the trees! Throw up a handful of
feathers, and all must fall to the ground according to definite laws; but how
simple is this problem compared to the action and reaction of the innumerable
plants and animals which have determined, in the course of centuries, the
proportional numbers and kinds of trees now growing on the old Indian ruins! The
dependency of one organic being on another, as of a parasite on its prey, lies
generally between beings remote in the scale of nature. This is often the case
with those which may strictly be said to struggle with each other for
existence, as in the case of locusts and grass-feeding quadrupeds. But the struggle
almost invariably will be most severe between the individuals of the same
species, for they frequent the same districts, require the same food, and are
exposed to the same dangers. In the case of varieties of the same species, the
struggle will generally be almost equally severe, and we sometimes see the
contest soon decided: for instance, if several varieties of wheat be sown
together, and the mixed seed be resown, some of the varieties which best suit
the soil or climate, or are naturally the most fertile, will beat the others
and so yield more seed, and will consequently in a few years quite supplant the
other varieties. To keep up a mixed stock of even such extremely close
varieties as the variously coloured sweet-peas, they must be each year harvested
separately, and the seed then mixed in due proportion, otherwise the weaker
kinds will steadily decrease in numbers and disappear. So again with the
varieties of sheep: it has been asserted that certain mountain-varieties will
starve out other mountain-varieties, so that they cannot be kept together. The
same result has followed from keeping together different varieties of the
medicinal leech. It may even be doubted whether the varieties of any one of our
domestic plants or animals have so exactly the same strength, habits, and
constitution, that the original proportions of a mixed stock could be kept up
for half a dozen generations, if they were allowed to struggle together, like
beings in a state of nature, and if the seed or young were not annually sorted. As species
of the same genus have usually, though by no means invariably, some similarity
in habits and constitution, and always in structure, the struggle will
generally be more severe between species of the same genus, when they come into
competition with each other, than between species of distinct genera. We see
this in the recent extension over parts of the United States of one species of
swallow having caused the decrease of another species. The recent increase of
the missel-thrush in parts of Scotland has caused the decrease of the
song-thrush. How frequently we hear of one species of rat taking the place of
another species under the most different climates! In Russia the small Asiatic
cockroach has everywhere driven before it its great congener. One species of
charlock will supplant another, and so in other cases. We can dimly see why the
competition should be most severe between allied forms, which fill nearly the
same place in the economy of nature; but probably in no one case could we
precisely say why one species has been victorious over another in the great
battle of life. A
corollary of the highest importance may be deduced from the foregoing remarks,
namely, that the structure of every organic being is related, in the most
essential yet often hidden manner, to that of all other organic beings, with
which it comes into competition for food or residence, or from which it has to
escape, or on which it preys. This is obvious in the structure of the teeth and
talons of the tiger; and in that of the legs and claws of the parasite which
clings to the hair on the tiger’s body. But in the beautifully plumed seed of
the dandelion, and in the flattened and fringed legs of the water-beetle, the
relation seems at first confined to the elements of air and water. Yet the
advantage of plumed seeds no doubt stands in the closest relation to the land
being already thickly clothed by other plants; so that the seeds may be widely
distributed and fall on unoccupied ground. In the water-beetle, the structure
of its legs, so well adapted for diving, allows it to compete with other
aquatic insects, to hunt for its own prey, and to escape serving as prey to
other animals. The store of nutriment laid up within the seeds of many plants
seems at first sight to have no sort of relation to other plants. But from the
strong growth of young plants produced from such seeds (as peas and beans),
when sown in the midst of long grass, I suspect that the chief use of the
nutriment in the seed is to favour the growth of the young seedling, whilst
struggling with other plants growing vigorously all around. Look at a
plant in the midst of its range, why does it not double or quadruple its
numbers? We know that it can perfectly well withstand a little more heat or
cold, dampness or dryness, for elsewhere it ranges into slightly hotter or
colder, damper or drier districts. In this case we can clearly see that if we
wished in imagination to give the plant the power of increasing in number, we
should have to give it some advantage over its competitors, or over the animals
which preyed on it. On the confines of its geographical range, a change of
constitution with respect to climate would clearly be an advantage to our
plant; but we have reason to believe that only a few plants or animals range so
far, that they are destroyed by the rigour of the climate alone. Not until we
reach the extreme confines of life, in the arctic regions or on the borders of
an utter desert, will competition cease. The land may be extremely cold or dry,
yet there will be competition between some few species, or between the
individuals of the same species, for the warmest or dampest spots. Hence,
also, we can see that when a plant or animal is placed in a new country amongst
new competitors, though the climate may be exactly the same as in its former
home, yet the conditions of its life will generally be changed in an essential
manner. If we wished to increase its average numbers in its new home, we should
have to modify it in a different way to what we should have done in its native
country; for we should have to give it some advantage over a different set of
competitors or enemies. It is good
thus to try in our imagination to give any form some advantage over another.
Probably in no single instance should we know what to do, so as to succeed. It
will convince us of our ignorance on the mutual relations of all organic
beings; a conviction as necessary, as it seems to be difficult to acquire. All
that we can do, is to keep steadily in mind that each organic being is striving
to increase at a geometrical ratio; that each at some period of its life,
during some season of the year, during each generation or at intervals, has to
struggle for life, and to suffer great destruction. When we reflect on this
struggle, we may console ourselves with the full belief, that the war of nature
is not incessant, that no fear is felt, that death is generally prompt, and
that the vigorous, the healthy, and the happy survive and multiply. |